Tribe IX . RAMAL I N E I Nyl. Bull. Soo. Liiiu. Normand,
sor. 2, iv. (1870) p. 103.
Thallus fruticuloso, laciniose, or filamentose, rounded or com-
qiressod, erect or pendulous, in ternally with woolly arachnoid medulla.
Apotheoia leoanorine, soutellate, terminal or la te ra l ; spores
8næ, 1-septate, snboblong, stra ig h t or slightly curved, colourless ;
paraphyses discrete. Spermogones immersed or slightly prominent ;
sterigmata suhsimple or pauoi-articulatc, with loug anastomosing
filaments intermixed.
This tribe, under which Nylander in his former arrangement included
Alectoria and Erernia, is now, in its more limited acceptation, regarded
by him as quite distinct, on account of marked analytical differences,
more especially of the tbalamiuni, while tbe structure of the spermogones
is entirely peculiar (vide Kamal. ilouogr. I, c.). Tbe exotic genua liamalea
Nyl. externally resembling Eamalina, ought, from the type of the spermatia,
to be relegated to the Cladoniei ; while Dactylina and Eiifourea,
neither of which occur in Britain, are also excluded fro.u this tribe.
36. EAMALINA
Aoh. Lich.Univ. (1810)
p. 122.—Thallus coes-
pitoso-frutioulose, foliáceo
- complánate or
rounded, somewhat
shining or subopaque,
soft or rigid, solid or
fistulöse, raraoso-laoini-
ate,concolorous on both
sides ; medullary layer
arachnoid, becoming
sometimes very lax, cortical
layer composed
e ith e r of indistinct cells
or of longitudinal cong
lutínate filaments.
xApothecia scattered,
oqiaqne, subooncolorous
with thallus ; hq'pothe-
eium colourless ; spores
small ; paraphj’ses th in
or moderate, thicker or
clavate a t the apices ;
hymenial gelatine b lu ish,
th en violet with
Fig. 39.
Eamalina fraxinea Acb. —c. Longitudinal section
of (one side of) thallus, X200. h. Theca and
paraphysis, X 350. c. Spores, X5Ü0. d. Sections
of two spermogones, x30. e. Sterigmata
and spermatia, X 500. (Also on the left a
fragment of the anastomosing spermogonie iila-
nients.)
iodine. Spermogones sca ttered ; spermatia straight, cylindrical, or
oblongo-cylindrioal.
The species of this genus, formerly little understood, liave recently had
new light thrown upon them by the minute researches of Nylander. He
has shown th at the chemical reactions of the medulla with K, the differences
in the receptacle of the apothecia, the form and size of the spores
and spermatia, afford invaluable aid in the discrimination of species. The
number of species liave been consequently largely increased, and varieties
more definitely referred to their species. As compared with other European
countries, the genus has a very fair number of species and varieties
in Great Britain, all belonging to Nylander’s Section C, which is characterized
by tbe spermogones having pale or colourless conceptacles.
a. Thallus a tten u a te , fruticulose, subrounded or suhcompressed ;
cortical layer filamentose.
1. R. th r a u s ta Nyl. Bull. Soc. Linn. Normand, ser. 2, iv. (1870)
p. 116.—Thallus elongate, pendulous or prostrate, filiform, sub-
rounded, here and there oomqiressed, very much branched, smooth,
somewhat shining, pale straw-coloured ; laciniæ very slender, densely
interwoven, the apices unequally caqiillari-attenuate (medulla K—).
“ Apothecia superficial, sessile, minute, qilane, pale, th e margin very
thin; spores s tra ig h t.”— Cromb. Journ. Bot. 1872, p. 71 ; Leight.
Lich. Fl. ed. 2, p. 470, ed. 3, p. 83.—E amalina calicaris,vaT. thrausta
Cromb. Lich. Brit. p. 25 pro pa rte ; Leight. Lich. Fl. p. 94 pro
parte. Alectoria thrausta Ach. Lioh. Univ. (1810) p. 596.
The tballus, wbicb is somewhat Alectorioid, is very fragile when dry,
and in the few British specimens gathered is destitute of the soredia,
with which it is elsewliere sometimes sprinkled. The apothecia, 'which
are not well known, are absent from our specimens, which are also without
spermogones.
Hah. On sandy soil among short beath in maritime tracts.—Histr.
Very sparingly in one spot on the N.E. coast of Scotland.—B. M. : Bay
of Nigg, Kincardineshire (now extinct).
h. Thallus comjiressed, longitudinally striato-nerved or subcostato-
unequal ; cortical lay er filamentose.
2. E. calicaris Nyl. Bull. Soc. Linn. Normand, ser. 2, iv. (1 8 7 0 )
p. 131.— Thallus subrigid, compressed, lin e a r or lineari-laciniate,
pale glaucous or greyish green, laciniæ sublacunosely longitudinally
nervoso-rugose, usually canaliculate (meduUa K —). Apotheoia
pedicellate, marginal and terminal, small or moderate, pale fiesh-
coloured or glaucescent, th e receptacle rugose beneath ; spores
straight, ellipsoid, 0 ,0 1 0 -1 6 mm. long, 0 ,0 0 5 -7 mm. thiok.—Cromb.
Grevillea, xv. p. 47 ; Lich. Brit. qi. 25 pro q>arte ; Leight. Lioh. FL
ed. 3, p. 83.—E am a lin a calicaris y . canaliculata Fr. llu d d , Man.,
p. 73, t. 1. f. 17 ; Leight. Lich. F l. p. 92. Eamalina fa stigiata [3..
calicaris Hook. Fl. Scot. ii. p. 68 pro p a rte ; Tayl. in Mack. Fl. Hib.
ii. p. 85. Loharia calicaris Hoffm. Deutsch. El. ii. (1795) p. 139
pro parte. Lichen calicaris Huds. Fl. Augl. p. 451 pro parte ; Lightf.
Fl. Soot.ii. p. 834 qiro p a rte ; With. Arr. ed. 3, iv. p. 51 pro parto.
Lichen fa stigiatus Eng. Bot. t. 890 (upper fig.). Eamalina fa stig ia ta
Sm. Eng. Fl. v. p. 225 pro p arte. Lichenoides coralliforme rostraium et
canalieulatum Dill. Muse. 170, t. 23. f. 62 a.—Lichenoides arboreum
lil
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