1-2 inches long, ovate, simply serrate with broad mucronate teeth. Cymes
shorter than the leaves: bracts subulate. Calyx persistent, cleft rather
below the middle ; the segments mostly acute, two of them somewhat narrower
; one of the broader ones sometimes minutely 3-toothed at the apex.
Petals twice or more the length of the calyx. Stamens deciduous. Styles
more than twice the length of the ovary, much exserted beyond the calyx.
Ovary free, except the base, the parieties rather thick and firm; the dissepiments
very short; the placentae lunate, at first distinct, many-ovuled.—
We much regret that we have not more adequate materials for describing
this plant. Our specimens were collected by Dr. Edwin James (in Long’s
Expedition), but the particular locality is not recorded. It is probably rare
or very local, as no other botanist seems to have met with it. It appears to
be an entirely distinct genus, to which we have applied the present name in
commemoration of the scientific services of its worthy discoverer, the botanist
and historian of ‘ Major Long’s Expedition to the Rocky Mountains, in
the year 1820,’ and who, during that journey, made an excellent collection of
plants under the most unfavorable circumstances.
S uborder IV. PHILADELPHEE.
Ord. Philadelphe®, Don, DC. (excl. gen.)
Estivation of the petals convolute. Capsule opening by locu-
licidal dehiscence.—Shrubs, with opposite and simple exstipulate
leaves.
17. PHILADELPHUS. Linn. ; Lam. ill. t. 420 ; Gcertn.fr. t. 35.
Tube of the calyx obovate-turbinate, adherent to the ovary; the limb
4-5-parted, persistent. Petals 4-5, broadly obovate, convolute in aestivation.
Stamens 20-40, shorter than the petals: filaments filiform. Styles usually'
4, more or less united, sometimes nearly to the summit: stigmas oblong or
linear. Capsule mostly 4-celled, free at the summit, 4-valved, loculicidal;
the placentae projecting into the cells, many-seeded. Seeds pendulous and
densely imbricated downwards on the thickened placentae, scobiform; the
testa membranous and loose, subulate-attenuate at the apex, and with a short
lacerate appendage next the hilum. Embryo nearly the length of the thin
fleshy albumen: radicle cylindrical, much longer than the oval flatfish
cotyledons.— Shrubs (natives of North America and Japan ? or Central
Asia?), with opposite often serrate exstipulate leaves, and large racemose-
cymose or solitary (white) flowers.
1. P. inodorus (Linn.): glabrous; leaves ovate or ovate-oblong, obtuse at
the base, acute or somewhat acuminate, triplinerved, entire or very nearly
s o ; flowers (rather small) 1-3 at the extremity of the branches; segments of
the calyx triangular-ovate, acute, about the length of the tube ; style longer
than the stamens.—Linn. spec. 1. p. 470 (Catesb. Car. 2. t. 84); Walt. Car.
p. 146; Willd. spec. 2. p. 948; Bot. mag. t. 1478.
Carolina to Alabama, apparently confined to the upper country and somewhat
rare. Near Milledgeville, Georgia, Dr. Boykin! Upper part of
Alabama, Mr. Buckley ! May..—Whole plant glabrous. Flowers scentless.—
This species appears to be little known in cultivation. The flowers
are smaller and much less showy than P. grandiflorus.
2. P . grandiflorus (Willd.): more or less pubescent; leaves ovate or
ovate-oblong, acuminate, dentate or denticulate with sharp teeth, triplinerved;
flowers (large) 1-3 or more at the extremity of the branches, on slender
pedicels; segments of the calyx ovate or ovate-lanceolate, conspicuously
acuminate, much longer than the tube ; style equalling or longer than the
stamens.— Willd.! enum. 1. p. 5 1 1 ; Pursh! fl. 1. p. 329; £11. sk. 1. p.
538; Guimp. Otto, 8f Hayne, holz. t. 44 ; Schrad.! in DC. prodr. 3. p. 206,
Sc in Linntsa, 12. p. 43. P . inodorus, Michx.! fl. 1. p. 283. P. speciosus,
Schrad.! 1. c.
0. laxus: branches weak and pendulous ; leaves (especially of the young
branches) sharply toothed, the upper ones acute at the base and often entire;
segments of the calyx much elongated and spreading in fruit ; flowers 1-3,
or often 6.—P. laxus, Schrad.! 1. c .; Lindl. ! hot. reg. (ser. 2) t. 39.
y. floribundus: flowers 5-7.— P. floribundus, verrucosus, latifolius, &
Zeyheri ? Schrad. 1. c. P. grandiflorus, Bot. reg. t. 570.
Along streams, Virginia! to Georgia! mostly in the upper country. April-
May.—Shrub 6-10 feet high; the young branches long and flexible. Flowers
nearly inodorous, showy, usually only 3 together in a wild state.—The
several species of Schrader cited above, are certainly only varieties of our
P. grandiflorus.
3. P . hirsutus (Nutt.): leaves ovate, acuminate, sharply serrate-dentate,
3-nerved from the base, scabrous-pubescent above, canescently hirsute with
appressed hairs beneath, as well as the pedicels and calyx ; flowers (small)
1-3, terminating the short branchlets, on very short pedicels; segments of
the calyx triangular-ovate, about the length of the tube; style shorter than
the stamens; stigmas short, connate.—Nutt.! gen. 1. p. 301; D C .! prodr. 3.
p. 206. P. trinervius, Schrad., in Linncea, 12. p. 47. P. pubescens, Bose.
Tennessee, “ on the rocky banks of French Broad River, near the Warm
Springs,” Nuttall!—A small shrub, with virgate branches; the flowers in a
wild state smaller than in P. eoronarius. Styles connate to the summit, but
when old separable for one-third their length.
4. P . Leunsii (Pursh): leaves ovate, acute, 3-5-nerved from the base, the
adult ones nearly entire and somewhat glabrous; flowers (small) racemose,
on short pedicels ; segments of the calyx ovate-lanceolate, acute, twice the
length of the tube ; style about the length of the stamens; stigmas 3-4, very
long.—Pursh! fl. 1. p. 329 ; Hook. ! fl. Bor.-Am. 1. p. 220 (partly).
Oregon, Lewis! Douglas! Nuttall ! Mr. Tolmie ! in open pine woods.
Also near St. Barbara, California, fide Nuttall.—A low shrub, with slender
branches spreading horizontally, somewhat hairy when young. Leaves
small, the younger ones toothed; the adult ones often with a woolly tuft at
the axils of the nerves beneath. Flowers much smaller than in P. inodorus,
scentless. Fruit turbinate, 4-celled; but one of the stigmas often abortive.
5- P • Gordonianus (Lindl.): leaves ovate, acuminate, triplinerved, serrate
toothed, hairy ; flowers (rather large, numerous) in 5-9-flowered compact
racemes; segments of the calyx ovate, acuminate; style shorter than
the stamens, deeply-cleft; stigmas short.—Lindl.! bot. reg. (misc. 1838. no-
23, A) ser- '2- t. 32. P , Oreganus, Nutt. 1 mss.
Shady woods, Oregon, common near the coast, Douglas ! Nuttall! Mr.
Tolmie ! July.—Shrub 4-6 feet high. Flowers inodorous. Fruit large,
more than half superior. Leaves often very coarsely serrate and more or
less hirsute.