examination of the microscopic preparations lias been made. The form agrees with the
typical British examples. Malmgren’s artist had omitted the cilia on the ventral cirrus,
while in the description they are not mentioned. Baron de St.-Joseph, in a recent
publication,1 alludes to the same form, and appears to agree with the original author in
thinking it necessary to make a variety.
Another well-marked variety was procured between tide-marks, Herm, under a stone
in August, 1868, and also at Lochmaddy, North Uist. While in regard to the general
outline, the shape of the head, the position of the eyes, and the arrangement of the
pigment on the dorsum, it agrees with the ordinary form, the dorsal bristles are shorter,
though they do not deviate from the typical structure; and the same may be said of the
ventral. The scales, however, vary, since the cilia on the outer border-are shorter than
usual, and have large ovate heads containing granular epidermic elements; some of these
also occur within the posterior border, while along the whole of the latter is placed a
short series of cilia with large globular heads and short stalks, the terminal ones only
towards the inner border becoming ovoid instead of globular. The general surface of
the scale is studded with larger,tubercles ; and few of the others, except towards the inner
region, and one or two of those near the posterior border, assumed considerable proportions,
while none were so large as usual. A similar variety in regard to scales was
procured by Dr. G-wyn Jeffreys in 1868 in ninety fathoms off North TJnst, Shetland, but
the ventral bristles were more slender, both somewhat approaching those of E. Johnstoni,
though the dorsal were more tapered towards the tip. The specimen was a male.
2. E varne J ohnstoni, McIntosh, 1876.
Specific Characters.—Distinguished from E. impair by the deep brownish hue of the
dorsum, and the brownish-purple proboscis; eyes more minute, the anterior pair not
visible from the dorsum. Scales with fewer and larger horny papillae, longer, fewer,
and more delicate cilia along the- posterior and outer borders. Dorsal bristles more
slender and less tapered distally; ventral more slender, and with longer bifid tips.
S ynonym.
1876. Evarne Johnstoni, McIntosh. Trans. Z. S., ix, p*. 398, pi. vii, f. 13—18.
Habitat.—Dredged at Station 3, ‘ Porcupine,* 1870, in 690 fathoms in the Atlantic;
Station 115, West Ireland, August 20th, 1890 (Mus. R. Coll. Science, Dub.).. , Ranges to
Norway.1 2
Length about 9 mm.
1 'Ann. d. sc. n at./ viii séi’., v, p. 231, 1898.
2 Canon Norman, 1879, Stat. 30—34, 41 and 44.
The head (Plate XXVII, fig. 7, from a large example, and fig. 12, small example),
has the same form as in E. impar, and is pale throughout in the preparations, the absence
of pigment at the base of the tentacle being noteworthy. The eyes appear only as
minute black points; two lie at the posterior border of thé head, almost hidden by the
collar; two laterally in front of these, as in the ordinary species. A variety dredged
by the ‘ Porcupine ’ in 1870 at 690 fathoms has very large eyes (Plate XXVII, fig. 7),
the anterior pair having a lens-like corneal thickening. Adult Norwegian examples
show still further increase in these organs.
Body apparently similar in shape to E. impar, viz. abruptly diminished in front and
gently narrowed from behind.the anterior third. The dorsum has a deep brownish hue,
with a tinge of purple in front—from the proboscis. Behind the latter the pigment is cut
into bars by. the pale belt at the junction of each segment. The ventral surface is pale,
the margins of the oral aperture alone being deeply tinged with brown. The segmental
eminence is distinct, but the papilla is minute.
In the examples the proboscis is more or less extruded, and the usual number of
conical papillse (nine) fringe each lip.
Scales.—A single reniform scale occurred in the vessel, and from what is observed
in the examples from Norway it would seem to belong to the specimen. The surface,
with the exception of the inner fifth, has rather large conical horny papillae, often with
blunt spinous tips, while the adjoining external and part of the posterior border have a
few long and very slender cilia, with a somewhat fusiform tip. The contrast, therefore,
with the more densely and minutely spinous scale of E. impar is marked; the cilia,
moreover, on the outer edge of the scale of the latter are more numerous, larger and
longer, and globose at the tip.
Feet.—As in E. impar, the second foot presents shorter dorsal bristles with less
tapered tips, but otherwise they are of similar structure to the succeeding. The ventral
bristles of this foot do not project more than the dorsal, and hence are short as well as
slender, the long attenuated spinous region ending in a hair-like tip.
In the typical foot (Plate XXX, fig. 6) the dorsal division bears longer and more
slender bristles than in E. impar, the slight tapering towards the tip being a noteworthy
feature, and the rows of spines are even more distinctly marked. The smooth terminal
portion is decidedly shorter than in the latter, and in some cases it presents a slight mucro
' at the extremity, then a shallow notch, and another elevation a little above the first row
of spines. A long clear shaft projects beyond the foot before the rows of spikes appear,
so that the bristles are comparatively long. One of the stronger bristles is represented
in Plate XXXIX, fig. 23, while the tendency to differentiation of the tip is observed in
fig. 24, Plate XXXIX. The superior ventral bristles have tips so attenuate that it is
difficult to make out their structure; but the bifid condition is present, with the
exception perhaps of the first. The next series have much longer and stronger distal
regions, with extremely delicate and translucent bifid tips; the terminal hook is short
and very slightly curved, and the secondary process is rather short and broad, and passes
far up, while the rows of spines are distant and well marked. The tips of the succeeding
(lower) bristles become broader and shorter, but the character of the termination
remains the same. Toward the inferior edge the tip is simple, only a faintly developed
47