of elongated papillae, which are conical and taper to a point. They are best developed towards
the tip of the organ. Prof. Bourne describes each as furnished with a sensory hair.
■Body.—The dorsum is marked in the median line by broad interrupted patches of
blackish-brown pigment, three or four of the most prominent, anterior touches having a
pale area in the centre. These touches extend outward to the feet posteriorly; moreover,
dorsally the bases of the feet have black pigment, which becomes strongly
pronounced after the anterior third, the effect of the mottling of the feet and body
posteriorly being somewhat like that on tortoise-shell.
The ventral surface has a similar arrangement of areas to that in L. squamatus, only
the lateral are marked off from the bases of the feet posteriorly by a slightly elevated and
pigmented border, which terminates at the bases of the caudal styles, and is connected
with touches which extend a short way forward in the median line.
Posteriorly the body terminates in a grooved pedicle for the large symmetrical
caudal styles, all these parts being better marked than in L. squamatus. The anus is
situated dorsally at the base of the pedicle, to which it sends a ridge-like process.
The bristled segments are twenty-six.
Proboscis aiid Digestive System.-—De Saint-Joseph found sixteen papillas along the
margin of the proboscis, but there are eighteen, as in L. squamatus, in all the examples
examined. They are speckled with black pigment. In transverse section the organ is
typical.
Only two cylindrical and rather massive caaca pass forwards into the peripharyngeal
space, and their extremities appear to be devoid of any distinct differentiation. The
stomachal region of the gut anteriorly is remarkably muscular, the fibres at the front end
forming a row of separate dull orange lobes which clasp the proboscis, and which resemble
a series of glands. Their tissue, however, is wholly muscular, thepowerful fibres forming
loops in the rounded lobes, which are sometimes made by rupture of the fibres from
the proboscis.
Colour.—The dorsum is of a dull brownish hue speckled with white,, and with dark
touches on the scales. The under surface is pale, with some dark touches at the mouth
and the sides of the tail. The segmental papilla has a dark column and a whitish tip in
the posterior half of the body. At the reproductive season the males are pale under
the scales, the females dark grey.
Scales.—The scales (Plate XXXII, fig. 2) of this species are more or less circular
throughout, and do not quite cover the dorsum, or, as Marenzeller says, leave bare
rhomboidal spaces. Some are firmly fixed, .others easily separate from the pedicle as in
L. squamatus. They are more flexible than in that species, and, with the exception of the
first four pairs, show only minute tubercles. The-first, second, and third have numerous
small tubercles or blunt dumb-bell shaped papillae, distributed generally over the surface,
while the fourth has a smaller number, less distinctly raised above the surface. The
edge of the scale throughout is smooth. They are fixed by the umbilicus, which is
nearer the outer than the inner margin, this area being generally marked dorsally by a
white patch bordered with black pigment. The scales give the dorsum generally a
leaden hue, or in some a dusky brown, speckled with white. Besides the white patch,
which posteriorly is sometimes reniform, at the umbilicus the scales are mottled with a
blackish or slightly glistening dull leaden hue, or with more numerous black specks,
which posteriorly do not seem to be connected with papillas. The under surface is
smooth and iridescent, and has the opaque whitish patch of the umbilicus. In spirit
the anterior scales generally show a fold from the umbilicus to the external margin.
The minute structure of the processes of the scales has been specially described and
figured by De Saint-Joseph,1 who states that the tip is “ scaled ” and spinous. His
figure, indeed, is regularly and closely diced. So far as can be observed in the British
specimens, it is the tip and neighbouring part of the capstan-like column of the process
of the scale which is thus covered, as recently described and figured.2
Feet.—On the dorsum of the first foot is a minute tuft of tapered serrate bristles
(Plate XXXYII, fig. 14), and a single large spine which penetrates the skin beneath
the former. Prof. Bourne speaks of four bristles being borne by each division. This
is probably a misprint, as his figure differs.
The second foot diverges from that of L. squamatus in having its comparatively large
ventral cirrus directed more distinctly inwards and forwards, in the smoother foot and
the more translucent bristles. Moreover, while the dorsal series more or less correspond
at first sight, yet the character of the serrations slightly differs from those of L. squamatus,,
being somewhat shorter and finer, and the tips of the bristles are more tapered.
The ventral bristles of L. clava again differ in having their long tips spinous to the apex,
whereas in L. squamatus the spinous tips are shorter, the tip is bare, and in some slightly
hooked.
In the third bristled foot (the second of some authors) the dorsal bristles are less
gradually tapered than in L. squamatus, while the ventral series have longer spinous
regions bare in the upper at the tip—which is more slender than in L. squamatus ; indeed,
the whole bristle is more slender than in the latter species, and the rows of spines longer.
Both dorsal and ventral cirri, moreover, are different, being proportionally larger and
more bulbous below the filiform extremity.
In the fully-formed foot (e. g. the tenth, Plate XLII, fig. 26) the papilla for the
dorsal bristles is more prominent than in L. squamatus, and has a distinct black bar in
front, and the bristles are somewhat shorter and more curved (Plate XXXYII, fig. 15).
The tips of the ventral bristles are shorter, the curve more pronounced (Plate XXXYII,
fig. 11), and the bare portion shorter. The stronger and larger hooked tip in L. squamatus
is a characteristic feature. The segmental papilla is visible on the eighth bristled
foot as in L. squamatus, and is continued to the last foot.
In the terminal foot the dorsal bristles are shorter and more curved than in L. squamatus,
a feature present throughout the body. The ventral bristles have the curvature of
the shorter tip more pronounced than in L. squamatus. In both species these bristles
preserve great strength.
A marked difference between the species is the diminished size of the body and last
feet in L. squamatus, contrasted with the comparatively large size of all the parts in this
region in L. clava.
1 Op. cit., Aug., 1898, p. 229, f. 4, &c.
2 f Ann. Nat. Hist.,5 1898, p. 108, pi. ii.