less quadrate anteriorly, with the inner edge rounded. The straight external edge has a
series of about sixteen pinnate processes, besides a few simple papillae posteriorly. Each
process has a granular stem, to which is attached a series of lanceolate granular lamellae
with a narrow papillary tip. In the typical foot the inferior lobe is shorter than in S.
Mathildae, broadly clavate, and with a single papilla on its tip. The bristles are attached
to the whole tip (i. e. superiorly, terminally, and interiorly), and are long and minutely
spinous. The ventral lobe is devoid of the superior papilla seen in 8. Mathildas,.and .has
a more prominent process for the spine. The compound bristles (with jointed tips
—below the forms with the short tapering spike) have the terminal portion of the shaft
covered with whorls of somewhat sparse spikes, which are more numerous than in
8. Mathildas, while the stouter bristles beneath have close rows of minute spines on the
same portion of the shaft. One or two below the spine-papilla have short tips of one or
two segments. The slender ventral series have only about two whorls of spikes on the
end of the shaft, and the long jointed tips have, as in the upper series, minute beaks.
The ventral cirrus is long, slender, and subulate, extending beyond the fleshy part of
the foot.
S ynonyms.
1869. Sthenelais dendrolepis? McIntosh, Trans. R. S. E., vol. xxv, p. 409, pi. xii, f. 12; pi. xv,
f. 4, 5.
1876. Sigalion Bushii, McIntosh. Trans. Zool. Soc., vol. ix, p. 391, pl. lxx, f. 14.
Hahita^^Dredged in 90 fathoms off North TJnst, Shetland, by Dr. Gwyn Jeffreys, in
1867.
Head (Plate XXIX, fig. 7) somewhat pear-shaped in' outline, broad in front and
narrow posteriorly, where it is only separated from the nuchal collar by a slight furrow.
The anterior border is smoothly rounded, and abuts on the conical first foot. I t is quite
pale, no eye-specks being visible in the preparation. In the example a small and somewhat
clavate papilla (tentacle) projected from the centre of the anterior border of the
head—a remarkable condition in Sigalion}
Body somewhat stouter than in Sigalion Mathildae, and in this respect resembling
Sthenelais boa, but having the same general shape both dorsally and ventrally. The
example is incomplete, but it would seem to reach a similar length to the species first
mentioned. The segmental eminences exist at the bases of the feet posteriorly, but no
distinct papilla is visible. The general colour of the dorsum is pale greyish.
Scales.—The first pair of scales are almost ovoid, only the outer border is broader
than the inner, and has eight pinnate processes besides a smooth projection in front,
and some isolated papillge posteriorly. Except at the latter part the margin is quite
smooth. At the bases of the processes, however, isolated papillae occur as in S.
Mathildae, one or two of which are attached to the central axis as in the species just 1
1 The head had been injured, so that the question might be raised as to whether this was not
one of the lateral tentacles pushed out of position. After careful examination, however, the view
above mentioned was held.
mentioned. No branchial process occurs on the papilla supporting this scale,—indeed, it
only appears on the papilla of the fourth scale on each side.
Passing backward the scales assume a somewhat quadrate outline, which in a typical
example anteriorly has the inner end rounded, while the external border has about
sixteen beautifully pinnate processes, besides a few simple papilla* at one end (Plate
XXXIV, fig. 15). The pinnate processes have a tree-like figure—more robust than in
S. Mathildae, and the pinnae, instead of being hyaline cylindrical processes, are lanceolate
granular lamellae with a narrowed papillary tip (Plate XXVI, fig. 9). The posterior
scales are longer transversely, and have fewer pinnate processes, but the structure is
essentially the same. The granular epiderm (hypoderm) of the curve between the processes
is denser, more opaque, and in this condition extends up the stalk to the first pinnae.
Feet.—The first pair project forward just beneath the head, and are bluntly conical
with a single spine, which passes forward and upward near the dorsal cirrus—a slender
and subulate organ, with a slightly dilated tip in the preparation. It springs from the
outer and upper lateral region of the foot. The ventral cirrus arises from the lower
part of the foot, somewhat internal ,to a vertical line from the former, to which it is
similar in outline. The tuft of slender simple bristles is directed forwards and upwards;
they are minutely spiked. The palpus is fused with the base of the foot inferiorly, and
is shorter proportionally in the. specimen than in S. Mathildae. I t is finely tapered
and quite smooth. Only a blunt papilla occurs in the region of the lateral tentacle.
The second foot agreed with that in the former species, presenting a well-marked
though short dorsal lobe, with a spine and a tuft of simple bristles, and a somewhat
trifoliate ventral lobe with a. series of bristles. These have long slender shafts with a
few spikes on the dilated distal end, and a long jointed and finely tapered distal region,
with a slight trace of a bifid (beak-like) tip.
In the typical foot (Plate XXXI, fig. 11) the superior lobe is shorter than in
S. Mathildae, and broadly clavate, the expansion of the tip in lateral view being greater
than in the former species. It bears a brush of simple minutely spinous bristles, which
spring from the superior and inferior borders as well as the tip. The stronger and
longer forms arise above the spine; the more slender occur for the most part below it, and
especially the inner group. The papilla at the tip of the superior lobe appears a short
distance behind the front. Three ciliated pads (ctenidia) exist on the superior border
between the branchial process and the tip. A stalked infusorian is present on the bristles.
The ventral division is devoid of the superior papilla observed in S. Mathildae, and
has a more prominent papilla' for the spine. The ventral bristles are characterised, in
contrast with S. Mathildae, by having rather longer terminal processes with bifid tips.
The superior series which adjoin the forms with the short tapering spinous region have
the terminal portion of the shaft covered with whorls of Somewhat sparse spikes
(Plate XLII, fig. 18), which are much more numerous than in S. Mathildae, while the
stouter bristles below have close rows of minute spikes on the same portion of the shaft
(Plate XLII, fig. 19, and in some the rows' are closer than in the -example figured). One
or two below the spine-papilla have only a single articulation in the terminal region,
while others have two, and the ends of the shafts in these are smooth, or in the former
with only traces of spinous rows. The slender ventral bristles have generally about two