CONSPECTUS OR ABRIDGED CHARACTERS OF THE GENERA OF THE TRIBE
CUCURBITEiE* OF SCHRADER.
§ 1.—Filaments 5, inserted on the throat o f the corolla;
anthers distinct or 3 -adelphous, anticous, straight; •fruit,
------ baccate, few seeded.
1. Coniandra (Schrad.) Corolla 5-partite: connectiva
conniving, oblong, conical: fruit beaked. South
Africa. ' — .
2. C yrtonema (Schrad.) Limb o f corolla 5-partite :
filaments 5, incurved, connectiva incrassated 3-adel-
phous; anthers fixed laterally under the apex. South
Africa.
r 2.—Filaments d i or tri-adelphous, inserted on the tube
o f the corolla; anthers lateral, straight, ‘2-3-adelphous.
3. Sicydium (Schlecht.) Corolla 5-petaled, petals
undivided : filaments 3-adelphous, dilated and incurved
at the apex; anthers without a beak. Mexico.
4. B ryonopsis (Am.) Corolla 5-partite, lobes obo-
vate entire, undulated: filaments 3-adelphous, inserted
on the throat, straight; anthers pointless: stigma fringed
: berry few seeded. East Indies—Courtallwm.
5. Achmandra (Arn.) Lobes of the corolla undivided
: filaments tri-adelphous very short; anthers anticous,
inserted along the margins of the connectivum,
linear oblong; connectivum prolonged into a short beak
beyond the anther: fruit baccate (always?) beaked.
E a st Indies. Bryonia epigasa, rostrata, deltoidea, and an
undescribed species from Malabar.
Obs.—Perhaps this and the two last genera might be
joined to Melothria.
6. M elothria (Linn.) Lobes o f the corolla undivided,
denticulated: filaments 3-adelphous; connectivum
pointless: fruit baccate, not beaked. America.
Obs.—Schrader notices an East Indian one, but that
may perhaps be an Achmandra.
7. C eratosanthes (Schrad.) Lobes o f the corolla
linear bifid: filaments 3-adelphous. America.
8. A n g uria (Linn.) g Lobes of the corolla entire:
stamens diadelphous: fruit somewhat 4-angled. America.
§ 3.—Filaments 3-adelphous inserted on the top o f the
tube, anthers all cohering by means o f their connectiva,
and applied at the back along the margins o f the connectiva,
sigmoid. (?) two-celled.
9. Schizostigma (Arn.) Style simple, stigma peltate,
fleshy, cleft into 10-12 radiating linear lobes. Andes
o f Mendosa. ( Cucurbita asperata. Gill.)
§ 4.—Filaments distinct or tri-adelphous. inserted on the
throat o f the corolla, anthers 6 or 3-adelphous, gyrose, anticous.
10. S ph enanthe (Schrad.) Mexico.
k 5.—Filaments tri-adelphous, inserted at the base o f
the corolla, anthers lateral, straight, 3-adelphous.
11. P ilogyne (Schrad.) Calyx campanulate; segments
of the corolla patent, much longer than the calyx:
anthers one-celled: style entire; stigma 1, pileate: fruit
baccate, few seeded, obtuse. South Africa. [E a s t Indies—
Bryonia Garcini ?]
12. Z ehncria (Endl. ?). Lobes of the corolla quite
entire : style trifid; stigmas 3, flabelliform, quite entire:
fruit baccate, few seeded, blunt. South Africa, East
Indies and ? Norfolk Island.—Bryonia Mysorenses.
B . Hookeriana.
Perhaps this is only a subgenus of Pilogyne.
13. Kariv ia (Arn.) Calyx urceolate, corolla scarcely
exserted, lobes minute, quite entire ; anthers 2-cell-
e d ; style entire ; stigma pileate, 3-fid : fruit a peponida,
many seeded, blunt or with a short thick beak. East
Indies, Bryonia umbellata. B . amplexicaulis.
14. R hynchocarpa (Schrad.) Lobes of the corolla
denticulate, ciliated; style trifid ; stigmas 3, jagged and
to o th ed : fruit with a long slender beak. Guinea.
§ 6.—-Filaments tri-adelphous, inserted at the base o f
the corolla, anthers all cohering, posticous, linear, straight.
15. Must a (Arn.) Style entire, stigmas 3, more or
less cohering, erect. Anthers distinct, one-celled, lateral
; [connectivum prolonged, forming a projecting point:
a globose abortive ovary in the bottom o f the calyx.]
E a st Indies—Bryonia scabrella. \_B. leiosperma.]
§ 7.—Filaments 5, or tri-adelphous, inserted at the base
o f the corolla', connectiva toothed or lobed, anthers applied
at the back olong the margins o f the connectivum, and
therefore flexuose, gyrose, or anfractuose.
16. Bryonia (Linn.) Corolla 5-cleft: anthers tri-
adelphous, one-celled : style trifid; stigmas subreniform
or bifid: fruit ovoid or globose, baccate; few seeded.
Europe and East Indies—B.laciniosa, India. B . alba.
B . dioica, Europe. Perhaps also B . Garcini and leios-
perma, but o f these 1 have no male flowers by me to examine
[J5. Garcini. Stamens tri-adelphous one-celled:
cells linear, marginal, not sigmoid, style one, stigma dilated,
peltate : ovary few seeded ; pepo inverse reniform,
2 seeded. This is probably a new genus, but if not seems
to belong to Pilogyne rather than any other here. It
cannot possibly belong to Bryonia.’]
In the European plants, the type of the genus, there
are two ovules in each o f the 3 cells of the ovary—nearly
all Blame’s species belong to other genera.
17- C it r u l l u s . (Schrad.) Corolla persistent, 5-parted,
subrotate: anthers tri-adelphous, bilocular: style 3-fid;
stigma obcordate, convex: fruit a fleshy or dry and
fibrous, many-seeded peponida. Africa, East Indies.
Cucurbita citrullus and Cucumis colorynthis.
18. E cbaliam (Rich.) Corolla 5 c le ft: anthers tri-
adelphous : ovules in two rows in each c e ll; stigmas
three, two horned: fruit an elastically and irregularly
bursting peponida. Europe—Momordica elaterivm.
19. Momordica (Linn.) Petals 5, adnate to the base
of the calyx, deciduous: anthers all cohering: ovules
* This tribe contains all th e East Indian genera except Zanonica—Schrader refers Enythropapm, of Blume to
Cucurbiteae but that genus is very closely allied to Mackaya (Arnott in Jardine’s Mag. of*Zool. a nd Bot. vol. 2)#
and does not belong to the order.
in a single* row in each c e ll; stigma two lobed ; fruit a
capsular, elastically bursting, 3 valved peponida. E .
Indies, South Africa and America.
r* This though practically correct is not theoretically
so the carpellary structure being the same here as in
others; each margin has its placenta and ovules: and
though at any one section only one row appears, we do
not find the ovules always attached to the same line of
placenta on slicing the ovary successively from end to end
but sometimes on the one, sometimes on the other side of
the c e ll; such at least I find it in Momordica Charantia.]
To this genus seems to belong, Muricia, Loureiro and
Neurospermum, Raf.
20. Luffa (Cav.) Petals 5, inserted in the base of
the calyx, deciduous : anthers all distinct or di-triadel ■
phous : style 3 fid; stigma reniform or bipartite : fruit a
peponida, at length dry and internally fibrous, usually
opening by a terminal lid, rarely indehiscent. East Indies
and Arabia. There are 3 sections of this genus.
1st. Stamens distinct, Luffapentandra, acutangula, and
Kleinii.
2d. Stamens 3-adelphous. L. amara, Roxb. and nearly
all the species of Turia, Forsk.
3d. Stamens di-adelphous—L. tuberosa, Roxb.
21. Bbnincasa (Sav.) Corolla (yellow), five-parted,
patent: anthers 3-adelphous: style undivided, very
short; stigma large, thick, irregularly lobed and plaited :
peponida, fleshy indehiscent. Asia.
22. L agenaria (Ser.) Corolla (white) five petaled :
anthers 3-adelphous : style almost none; stigmas 3, thick
and 2-lobed: peponida fleshy and indehiscent. India,
South Africa.
§ 8.—Filaments, 3-adelphous, inserted on the tube o f
the corolla; connectiva entire, anthers 3 or mon-adelphous,
posticous, linear, bent upwards and downwards: calyx
long, tubular.
23. T richosanthes (Linn.) Segments o f the corolla
lacerated and fringed: anthers 3-adelphous ? or all united
: style trifid; stigmas oblong, subulate : fruit a peponida,
many seeded. E . Indies.
I reunite Involucrarea to this as a mere section depending
on the bracteas, the character taken from the
anther not holding good, at least T. Cucumerina has frequently
the anthers all united, and I suspect also T. an-
guina; perhaps they only become tri-adelphous after
[fecundating.
[In all the species I have had an opportunity of care-
; fully examining, the anthers are monadelphous or united.
The style is not trifid, nor properly speaking the stigmas
r subulate, as they cohere nearly to the apex by their cen-
| tral face, though the stigmatic surface extends for some
EXPLANATION
ZAN0NIA IND1CA.
[ 1* flowering branch (male plant) natural size.
I i ^ a*e flower, the petals removed, showing the 3-
| »bed calyx and insertion of the anthers.
I 3. An expanded flower showing the petals and stamens.
4. Corolla and calyx detached, the stamens more high •
[■ ? magnifted, to show the form o f the anther.
distance outwardly, and presents a somewhat subulate
outline.
In T. anguina they are never tri-adelphous, the anthers
cohere to the last as represented in the accompanying
figure. This last species with T. globosa, and tri-
foliata, Blume, and Involucrarea, Serange ( T . Wallich-
and) form a very characteristic section, perhaps a subgenus,
distinguished by their curiously bructeated male
flowers]. R. W.
24. Gymnopetalüm (Arn.) Calyx constricted at the
mouth; corolla (yellow) 5-parted; segments quite entire
: anthers all closely cohering: fruit baccate, ovate,
beaked, few-seeded: seeds large, roundish, with a blunt
margin E . Indies. There are two species—
1. G. Ceylanicum (Am.) Leaves deeply 5-lobed;
perianth glabrous: Bryonia tubiflora W. and A.
2. G. Wightii (Arn.) Leaves 3-5-angle-lobed; perianth
hairy. Courtallum.
§ 9.—Filaments usually tri-adelphous, inserted at the
base o f the perianth; connectiva entire, unless when produced
into appendages beyond the anthers: anthers linear,
posticous, bent upwards and downwards (calyx campanulate
or rarely infundibidiform.)
25. C ucumis (Linn.) Corolla 5-parted: anthers tri-
adelphous, or all of them slightly cohering, with appendages
at the apex! Peponida fleshy, indehiscent, or rarely
irregularly dehiscent, polyspermous: seeds ovate, compressed,
sharp edged. Asia, Africa and America.
26. Cucurbita (Linn.) Corolla campanulate, 5-cleft:
filaments tri-adelphous at the base or quite monadelphous
; anthers all cohering, without appendages: peponida
fleshy, indehiscent, polyspermous: seed with a
slightly thickened edge. Asia and America.
*27. Elaterium (Lin.) Petals scarcely united at the
base: filaments monadelphous, anthers all cohering;
style thick; stigma capitate : fruit a coriaceous, one-celled,
few seeded capsule, bursting elastically by two or
three valves. America.
28. S chizocarpum (Schlch.) Corolla infundibuli-
form, quite entire: filaments 3-adelphous: anthers all
cohering; peponida many seeded, bursting by several
valves that cohere by their apex. Mexico.
29. C occinea (W. and A.) Corolla campanulate,
segments acuminated: filaments monadelphous, anthers
tri-adelphous, conniving, without appendages : peponida
somewhat baccate, many-seeded. [Usually o f an oblong
oval shape and bright red when ripe.] E a st Indies.
§ 10.—Filaments monadelphous, connate into a column,
which is capitate at the apex, and then bearing the gyrose
posticous anthers.
30. Cephalandra (Schrad.) South Africa.
OF PLATE 103.
5. A fruit cut transversely.
6. A seed.
For this figure and dissection the draftsman is wholely
responsible. It was executed while I was in England,
and 1 have had no opportunity o f comparing it with living
plants to ascertain its accuracy. I suspect however
there is an error in making it a monoecious plant.