1911. AmpMcteis Gunnen, Ditlevsen. Danmark Exped., p. 427.
1912. „ ' „ Meyer. Inaug. Dissert. Kiel., p. 19.
» « » Wollebæk. Skrift. Vid.-selsk. Krist., No. 18, p. 55, pl. viii, figs. 1__11.
1914- „ „ McIntosh. Ann. Nat. Hist., ser. 8, vol. xiii, p. 98.
» » 3> Pauvel. Campag. Sc. Monaco, Fase. xlvi, p. 281, pl. xxvi, fig. 26.
” « « Ditlevsen. Polych. Annel. (Meddel. Grwfal,), p. 721.
1915. „ „ Allen. Journ. M. B. A., vol. x, p. 637.
1917. „ „ Hessle. .Zool. Bidrag, Uppsala, p. 116, Taf. i, fig. 9.
Habitat.—Stomach of the haddock, Sfc. Andrews Bay (E. M.) Dredged by Dr.
Gwyn Jeffreys off the Hebrides,» 6 6 , and in 80—100 fathoms in St. Magnus Bay,
Shetland; 90-125 fathoms, 30—50 miles West of Valentia, Ireland (J. G. J.) ;' “ Knight
Errant,” Stat. 8, 640 fathoms (yery large). Plymouth (Spenoe Bate and Rowe, Allen);
S.W. Ireland, log. 19, 75 fathoms, 1885 (A. C. H.) ; Kenmare, S.W. Ireland, 1886;
S.W. Ireland, log. 55, 1866, 23—38 fathoms (E. Wright). I. A.); Bay of Galway (E. p !
Abroad it has been found by the “ Challenger ” in 2750 fathoms in the mlOHle 0f the
Atlantic; Barents Sea (McIntosh); Naples (Claparède); Greenland (Grube); Norway
(Sars) ; Spitsbergen, Finmark (Norman). In 220 fathoms between Anticosti and the
south shore, Canada (Dr. Whheaves) ; Kerguelen (Ehlers)., New England, Ü.S.A.
(Verrill) and Atlantic Coast ; Kara Sea (Leyinsen) ; Arctic Sea (Marenzeller) ; Monaco
(Fauvel) ; West Indies (Augener).
The cephalic region (Plate CXVIII, fig. 7) is somewhat shield-shaped dorsally, with
a ridge running forward on each side anteriorly and a median groove. Each ridge
diverges a little and projects in front. A prominent fillet-ot the buccal segment bounds
them laterally, and converging to a median dimple guards them posteriorly as two
oblique ridges, whilst in front of these a less prominent V-shaped fillet lies immediately
behind the ridge marking the nuchal organs. On each side,At the commencement of the
posterior slope; is a minute eye—indistinct in most spirit preparations. A dimple in
the fillet opposite the eye-speck increases its range posteriorly, and represents the nuchal
organ with pigment-specks in front. The buccal segment has an irregular border
anteriorly, since, besides the two lateral fillets at the-cephalic plate, a narrow rim passes
in front of the mouth and a broader behind it, the margin of the lower lip being marked by
five crenations, the three median and their four grooves being most distinct. Projecting
from the mouth are the buccal tentacles, which are smooth. The second segment is
narrower than the foregoing, and has a nearly straight anterior margin dorsally, whilst
ventrally it is sinuous, having a forward curve in the middle, then a concavity, whilst the
lateral border again slightly curves forward. The region containing the foregoing parts
forms a blunt cone differentiated from the succeeding, which is wider, though the
maximum transverse diameter is four or five segments behind.
The branchiae are rather massive subulate organs springing from the third and fourth
segments and the anterior edge of the fifth. Each has a short basal region, from which
it readily separates, and a tapering distal part ending in a filiform tip. Two are anterior
and two posterior. In the smaller examples variation in the origin of the stems exists,
the outer anterior being sometimes nearly in a line with the origin of the posterior p at
of one side. In a large example, again, the outer anterior are external to a line from
the outer posterior.
The stomach of this form has a blind sac which opens into the ventral aspect of the
organ.B
ehind the sixth bristled segment the body gradually diminishes to the tail, which
terminates in a median anus with a lateral subulate cirrus on each side. The surface
is rounded and smooth dorsally, slightly flattened in front ventrally, and marked by
transverse glandular ridges, a distinct median groove running from the middle to the tip
of the tail. The length of the body varies from 1 to more than 2 inches.
The third segment, from its greater width and prominent anterior border, indicates
the commencement of the bristled region. Its dorsal margin is boldly concave forward,
whilst its ventral edge is nearly straight, and there is little to separate it ventrally from
the succeeding segment. It carries on the prominent lateral region the fan-like paleas,
which are more or less horizontal—that is, the concavity of the fan looks upward, the
convexity downward, and the longest bristles are internal, their number being from
fourteen to twenty. They are flattened golden bristles minutely striated longitudinally
(Plate CXXIV, fig. 5), the striae ending in granules distally, whilst the finely tapered tip
is translucent. A few transverse bars occur here and there on the shaft, which dilates
from the base to the surface of the skin, and then gradually tapers to the attenuate tip.
They form two groups, an anterior longer and a posterior shorter, the former probably
representing the dorsal division, the latter .the ventral.
The anterior region has seventeen pairs of dorsal bristles, the first two of which are
small, but the rest are conspicuous tufts projecting from setigerous processes which,
when viewed from above downward, are nearly cylindrical, but antero-posteriorly are
slightly tapered distally, and have at the ventral edge of the bristle-tuft a clavate papilla
which seems to have escaped Malmgren. This clavate papilla is less developed in front
than in the posterior setigerous processes, where it is much larger distally. The bristles
(Plate CXXIV, figs. 5 a, 5 a and 5 a") have straight striated shafts which dilate a little
from the base upward, continue of nearly equal diameter to the commencement of the
wings, and then curve slightly backward and taper to a fine tip. The strige of the shaft
become oblique in the curved terminal region, and the wings themselves are striated for
some distance upward. These bristles are evidently much used by the annelid, and the
basal striated portion of the wings is often worn. A transverse ridge with a small dorsal
cirrus—curved downward—represents the dorsal division behind the foregoing, and
continues to the tip of the tail.
The hatchet-shaped lamellae for the hooks commence on the ventral surface of the
seventh segment at some distance from the, setigerous process and at the posterior edge
of the segment. A more or less distinct ridge, wider anteriorly and gradually diminishing
posteriorly, connects them with the setigerous processes. The first are small and little
elevated, but they increase in prominence, and gradually approach the setigerous process,
so that at the twelfth or thirteenth bristle-bundle they are close to it; and the tip of the
last is nearly as prominent. Thereafter the uncinigerous processes form conspicuous
lamellas on each side of the posterior region to the tail. The uncinigerous lamella has in
the preparations a slightly irregular or creuulated edge to which the hooks are attached,
179